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Smithsonian Tropical Research Institute (STRI)

Joshua Tewksbury, Director

Thirty-five permanent scientific staff and twelve hundred visiting scientists each year at the Smithsonian Tropical Research Institute offer a dynamic intellectual community for students of tropical biology.  Our two areas of excellence: discovering and understanding life’s diversity, and the study of human diversity and cultural change, embrace a wide range of academic disciplines.  STRI is the world’s premier international research platform for tropical biologists working in more than forty countries.  We offer expertise, data, training programs and tools to young scientists and professionals alike. This enriches our own endeavor as STRI staff engage in collaborations and intellectual exchange with scholars and students from a wide range of partner institutions.

The Smithsonian’s history of 100 years of tropical research in Panamanian evolution, ecology and behavior of tropical organisms informs STRI’s innovative initiatives in soil science, plant physiology, canopy biology, paleoecology and climate change, molecular biology and neurobiology.

The Isthmus of Panama offers unique advantages for biodiversity research. Formed about three million years ago, the isthmus is a complex zone of contact between the floras and faunas of North and South America. In addition, it separates closely related forms of marine life that were isolated in the Atlantic and Pacific Basins. Panama’s topography and small size mean that a wide range of habitats are accessible within a short distance. Terrestrial habitats range from dry grasslands and scrub, through seasonal and evergreen lowland tropical forests, to montane and perpetually wet cloud forests. Marine environments include muddy, sandy, and rocky shores, coral reefs, mangroves, sea grasses and the open waters of the Caribbean and the eastern Pacific. Rivers and lakes provide opportunities for freshwater studies. Ten modern research facilities in the Republic of Panama provide an ideal base for tropical biology research.

STRI’s headquarters, including administrative offices, library, and the Earl S. Tupper Research and Conference Center, are located in the Ancon area of Panama City. The Tupper Center houses laboratories and offices for scientists as well as an auditorium and meeting rooms. STRI’s Tropical Sciences Library is one of the most comprehensive in the world for tropical studies, receiving 500 periodicals, containing more than 60,000 volumes and offering fast, online reference services. Barro Colorado Island (BCI) is one of the best known sites in the world for the study of lowland tropical forest. A reserve since 1923, research at BCI builds on almost 100 years of scientific studies that have resulted in over fifteen hundred publications. Facilities include modern laboratories and living quarters for approximately 65 visiting scientists. The 5,000-hectare Barro Colorado Nature Monument includes the island, five mainland peninsulas, and many smaller islands, which afford opportunities for the study of primary and secondary forests, freshwater habitats, and island biogeography.

STRI’s Center for Tropical Paleoecology and Archeology in Panama City is a five minute walk from its headquarters. This facility houses scientists, fellows and research assistants studying the geological rise of the Central American Isthmus, the evolution of tropical rainforests since the last glaciation, human colonization of the Isthmus and the origins of agriculture. STRI is developing major new laboratory facilities in the town of Gamboa intended to foster increased academic interchange among fields. The nearby22,000 hectare (54,362 acre) Soberania National Park,  contains lowland forest of various ages and a variety of freshwater habitats. A facility at Gamboa is also available for field courses. A station in Fortuna, Chiriqui, Panama’s western province, provides scientists with access to wet montane forests.

The STRI Tropical Forest Canopy Program uses construction cranes to study the upper canopy of tropical forests. Cranes are now in place at a dry tropical forest in the Parque Natural Metropolitano, in Panama City, and at a very wet tropical forest site on the Caribbean side, providing safe access to the forest canopies in contrasting environments. With marine laboratories on both the Pacific and Caribbean (Atlantic) coasts, STRI presents unique and unparalleled opportunities for comparative studies on the biota of the two oceans. Naos, on the Pacific, is a short distance from STRI headquarters near Panama City. The Molecular Evolution Laboratories, at the Naos Island Laboratories, host studies in evolutionary biology, genetics and molecular systematics. The Galeta Laboratories, 80 km (50 mi) north, on the Caribbean coast near the Atlantic entrance to the Panama Canal, is located at the edge of a fringing coral reef adjacent to mangrove forest. STRI’s Bocas del Toro Research Station is also located on the Caribbean on Isla Colón, near Panama's border with Costa Rica. This laboratory offers a range of services, including boats and SCUBA to provide investigators with access to an extraordinary diversity of marine and terrestrial biota.

STRI has expanded its research on animal behavior and evolutionary biology by establishing a laboratory of Evolutionary Neurobiology and Behavior. This facility focuses on comparative neurobiological and behavioral studies of tropical invertebrates with extreme body size reduction. The laboratory also provides well-equipped facilities for visiting neurobiologists; will host symposia on topics relating to brain miniaturization; and will explore connections between brain miniaturization and applied research related to engineering and information technology.

STRI scientists collaborate with research and academic organizations at sites throughout the tropics. STRI maintains formal cooperative research partnerships with colleagues in Kenya, Malaysia, India, Thailand, Sri Lanka, Ecuador, Brazil, Cameroon and other countries.

Large-scale, permanent forest plots at almost 80 sites in several dozen countries have been established based on standardized methods originally developed on Barro Colorado Island in Panama. These forest dynamic plots form a Global Earth Observatory network, ForestGEO, a STRI program to promote long-term biological and socio-economic research within tropical forests and forest-dependent communities, and to translate this information into relevant forest management, conservation, and natural resources policies. Many findings are tested directly by the Agua Salud Project in the Panama Canal watershed, a huge, landscape-level study of reforestation with native tree species and in close cooperation with the Panama Canal Authority and Panama’s Ministry of the Environment, MiAmbiente. At the Mpala Research Centre in the Laikipia Plateau of Kenya, scientists integrate research in disciplines ranging from wildlife/livestock management, ecology, geology, and animal physiology.

In addition to Smithsonian-wide fellowships administered by the Office of Fellowships in DC, STRI offers its own international, competitive awards to support the research of visiting students. Requests for fellowship information should be addressed to the Office of Academic Programs (Email: STRIFellows@si.edu). Information on facilities usage, housing and fees can be addressed to the Office for Visitors Services (Email: STRIRegistration@si.edu). Further information on STRI facilities and research programs can be found at STRI’s web site: https://stri.si.edu/

RESEARCH STAFF

Aiello, Annette, Staff Scientist. Entomologist. B.A. (1972), Brooklyn College, New York; Ph.D. (1978) Harvard University. Research specialties: Life histories, behavior, and evolution of insects, especially moths and butterflies. Contact: AielloA@si.edu

Amador, Sabrina, Staff Scientist. Animal Behavior , Evolutionary Biology , Natural History. B.S. (2004), University of Costa Rica; M. Sc. (2008) Universidad de Costa Rica; Ph. D.  (2014) University of Texas at Austin. Research specialties:  Behavioral ecology, functional morphology, ethology and evolution of behavior. Contact: amadors@si.edu

Broders, Kirk, Staff Scientist. Microbial Ecology , Disease Ecology , Evolutionary Biology. B.Sc. (2008) University of Nebraska–Lincoln; Ph.D., (2014) The Ohio State University. Population biology and evolutionary ecology of plant-associated microbes including pathogens, endophytes, saprophytes and mutualists. Contact:  brodersK@si.edu

Collin, Rachel, Staff Scientist, Director of the Bocas del Toro Research Station. Sc.B. (1993) Brown University; M.S. (1996) University of Washington; Ph.D. (2002) University of Chicago. Research specialties: Evolution of marine invertebrate life histories and development; systematics of marine gastropods. Contact: CollinR@si.edu

Cooke, Richard, Staff Scientist. Archaeologist. B.A. (1968) University of Bristol; Ph.D. (1972) University of London. Research specialties: Archaeology of New World tropics; faunal analysis; human ecology; history of fishing. Contact: CookeR@si.edu

Correa, Mireya D., Scientific Staff, Botanist. Licenciado (1963) University of Panama; M.A. (1967) Duke University. Research specialties: Tropical plant systematics, especially the flora of Panama; ethnobotany, especially medicinal plants. Contact: CorreaM@si.edu

Davies, Stuart, Senior Staff Scientist, B.Sc. (1987) University of Sydney, Australia; Ph.D. (1996) Harvard University. Research specialties: Tropical forest ecology; Asian ecosystems; plant ecology; long-term ecological observatories; plant taxonomy. Contact: StuartDavies@si.edu

Eberhard, William G., Emeritus. Biologist, Professor, Universidad de Costa Rica. A.B. (1965), Ph.D. (1969) Harvard University. Research specialties: Behavior of insects and web-building spiders; sexual selection by cryptic female choice; genitalic evolution, effects on behavior of miniaturization of nervous system. Contact: WilliamEberhard@si.edu

Guzman, Hector M., Staff Biologist. B.Sc. (1979), M.Sc. (1986) University of Costa Rica; Ph.D. (1994) Newcastle University, UK. Research specialties: Ecology and population dynamic of coral reefs; sclerochronology; conservation biology; human impacts on marine ecosystems, coastal management and marine pollution. Contact: GuzmanH@si.edu

Hall, Jefferson, Staff Scientist and Principal Investigator, Agua Salud Project. B.A. (1983) Miami University; M.F.S. (1992), Ph.D. (2002) Yale University. Research specialties: Tropical forest silviculture and management, restoration ecology, ecosystem services, secondary succession. Contact: HallJe@si.edu

Herre, E. Allen, Staff Scientist. Biologist. B.A. (1977) University of Chicago; Ph.D. (1988) University of Iowa. Research specialties: Coevolution of figs and their pollinator wasps; host-parasite coevolution; the evolutionary effects of population structure on sex ratio, sexual selection and parasite virulence; and interactions between tropical host plants and both mycorrhizal and endophytic fungi. Contact: HerreA@si.edu

Hubbell, Steven, Emeritus Staff Scientist. B.A. (1963), Carleton College; Ph.D. (1969) University of California, Berkeley. Research specialties: Tropical forest ecology; theoretical community ecology. Contact: StephenHubbell@si.edu

Jackson, Jeremy B.C., Emeritus. Biologist Emeritus. B.A. (1965), M.A. (1968) George Washington University; M.Phil. (1970), Ph.D. (1970) Yale University. Research specialties: Diversity, speciation and extinction of tropical marine invertebrates; ecology and conservation of tropical coastal communities.

Jaramillo, Carlos, Staff Scientist. B.A. (1992) Universidad Nacional de Colombia; Ms. (1995) University of Missouri-Rolla , Ph.D. (1999) University of Florida. Research specialties: palynology, paleobotany, paleontology, biostratigraphy, paleobiology. Contact: JaramilloC@si.edu

Kline, David, B. A. (1996), Ph.D. (2005). Scripps Institution of Oceanography, University of California, San Diego. Research specialties: Coral Reefs and Climate Change, Computer Vision Coral Ecology, Coral Reef Ecology and Coral Diseases, Cellular physiological mechanisms for coral calcification and photosynthesis- Extending Lab Based Models to the Field. Contact: dkline@ucsd.edu

Knowlton, Nancy, Emeritus. Sant Chair for Marine Science. A.B. (1971) Harvard University; Ph.D. (1978) University of California, Berkeley. Research specialties: Marine Biodiversity; ecology, behavior, evolution and systematics of coral reef organisms. Contact: Knowlton@si.edu

Larsen, Matthew, Director. BSc. (1976) Antioch College, Ph.D. (1997) University of Colorado. Research specialties: Tropical geomorphology and geomorphic work. Contact: LarsenMC@si.edu

Lessios, Harilaos, Senior Staff Scientist. B.A. (1973) Harvard College; M.Phil. (1976), Ph.D. (1979) Yale University. Research specialties: Molecular evolution and population genetics of marine organisms; ecology of coral reefs. Contact: LESSIOSH@si.edu  

McMillan, Owen, Staff Scientist. B.Sc. (1985) Duke University, M.Sc. (1991) University of Hawaii, Ph.D. (1994) University of Hawaii, Postdoctoral (1994-1997) University of College London. Research specialties: Evolution, behavior, origins and genetic basis of adaptive variation. Contact: McMillanO@si.edu  

Muller-Landau, Helene Clara, Staff Scientist. B.A. (1995) Swarthmore College; M.A. (1997), PhD (2001) Princeton University. Research specialties: Plant community ecology; Ecological and evolutionary theory; Anthropogenic influences on forests; Carbon budgets of forest ecosystems. Contact:  MullerH@si.edu  

O’Dea, Aaron, Staff Scientist. Paleobiologist. B.Sc. (1996) Liverpool John Moores University; Ph.D. (2000) University of Bristol. Research specialties: Tropical Paleobiology; Marine ecology and evolution; Historical Ecology; Conservation Paleobiology. Contact: ODeaA@si.edu  

Page, Rachel, Staff Scientist. B.A. (1996) Columbia University; Ph.D. (2008) University of Texas, Austin. Research specialties: Vertebrate behavior, sensory and cognitive ecology, predator-prey interactions, signal evolution. Contact: PageR@si.edu  

Piperno, Dolores R., Emeritus. Archaeologist/Paleobotanist Emeritus. B.A. (1971) Rutgers University; M.A. (1979), PhD (1983) Temple University. Research specialties: Archaeology and human ecology of tropical regions; paleobotany and paleoethnobotany. Contact: PIPERNOD@si.edu  

Robertson, D. Ross, Staff Scientist. B.Sc. (1966), Hons. (1968), Ph.D. (1974) University of Queensland, Australia. Research specialties: Zoogeography of neotropical shorefishes; Coral reef fishes: behavioral ecology; larval, reproductive and population biology; demography. Contact: drr@stri.org  

Roubik, David W., Staff Scientist. Entomologist. B.S. (1975) Oregon State University; Ph.D. (1979) University of Kansas. Research specialties: Tropical bees and beekeeping, ecology, systematics and behavior of bees, pollination systems; palynology; ecology and entomology. Contact: ROUBIKD@si.edu  

Salazar Allen, Noris, Scientific Staff, STRI, Research Professor and Curator of Bryophytes and Lichens, Herbarium, Department of Botany, Universidad de Panamá. B.A. (1969) Trinity College; M.A. (1973) State University of New York, Geneseo; Ph.D. (1986) University of Alberta, Edmonton. Research specialties: Biosystematics, phytogeography, morphology and evolution of tropical bryophytes; lichens of Panama. Contact: SalazarN@si.edu  

Saltonstall, Kristin, Staff Scientist. Molecular Ecologist.  B.A. (1992) Wellesley College; M.F.S. (1996) Yale School of Forestry and Environmental Studies; Ph.D. (2002) Yale University. Research specialties: Genetics of invasive species, invasive grasses, metagenomics, conservation genetics. Contact: SaltonstallK@si.edu  

Santos-Granero, Fernando, Staff Scientist. Social Anthropology. Licenciatura (1980) Pontificia Universidad Católica del Perú; M.Sc. (1981), Ph.D. (1986) London School of Economics and Political Science. Research specialties: Native Amazonian leadership and philosophies of power; history of native Amazonian peoples; Amerindian forms of slavery and servitude; tropical American chiefdoms; Amazonian regional economies. Contact: SantosF@si.edu   

Sharpe, Ashley, Staff Scientist. Archaeologist. B.A.. (2009) Boston University; Ph.D. (2016) University of Florida. Research specialties: Zooarchaeological Perspective on the Formation of Maya States. Contact: SharpeAE@si.edu

Slot, Martijn, Staff Scientist. Plant ecophysiologist. MSc (2003), Wageningen University, The Netherlands; Ph.D. (2013), University of Florida. Research specialties: Climate change impacts on photosynthesis, growth, and reproduction of tropical plants. Contact: SlotM@si.edu

Stallard, Robert, Staff Scientist. B.S. (1974), Ph.D. (1980) Massachusetts Institute of Technology-Woods Hole Oceanographic Institute. Research specialties: Earth-surface environmental processes with a focus on the study of water quantity and quality in streams, weathering, soil development, and erosion as related to land-cover and climate change in the tropics. Contact: Stallard@si.edu  

Torchin, Mark Erik, Staff Scientist. B.A. (1991), University of California, Santa Barbara; M.S. (1994) University of Oregon; Ph.D.(2002) University of California, Santa Barbara. Research specialties: Marine population and community ecology; host-parasite interactions; invasion ecology; conservation biology. Contact: TorchinM@si.edu  

Wcislo, William, Staff Scientist . Biologist. B.S. (1982) University of Michigan; Ph.D. (1991) University of Kansas. Research specialties: Evolutionary biology; ethology; behavior and behavioral evolution of bees, ants and wasps; perception and learning; social evolution; brain evolution. Contact: WcisloW@si.edu  

West-Eberhard, Mary Jane, Emeritus Staff Scientist. Entomologist. B.A. (1963), Ph.D. (1967) University of Michigan. Research specialties: Social behavior,natural history, and evolution of wasps (especially Vespidae); social evolution; sexual and social selection; developmental plasticity and evolution.

Windsor, Donald M., Emeritus Staff Scientist. Research Biologist. B.S. (1966) Purdue University; Ph.D. (1972) Cornell University. Research specialties: Macroevolutionary studies of Tortoise Beetle (Coleoptera, Chrysomelidae, Cassidinae) diversification related to food plant, parasitoid and mite associations worldwide. Contact: WINDSORD@si.edu   

Winter, Klaus, Staff Scientist. Biologist. Dr.rer.nat. (1975) Darmstadt; Habil. (1983) Würzburg. Research specialties: Plant Physiology; plant physiological ecology; photosynthesis and photoinhibition; crassulacean acid metabolism; plant responses to elevated CO2. Contact: Winterk@si.edu   

Wright, S. Joseph, Staff Scientist. Biologist. A.B. (1974) Princeton University; Ph.D. (1980) University of California, Los Angeles. Research specialties: Ecology of tropical forests; community ecology.

AFFILIATED RESEARCH STAFF

Angehr, George, Research Associate. BS (1973) Cornell University; Ph.D. (1980) University of Colorado. Research specialties: Ecology and distribution of the birds of Panama; Bird distribution in Latin America and Africa; Conservation of birds; Conservation of tropical forests. Contact: ANGEHRG@si.edu

Altieri, Andrew, Marine Biologist. B.A. (1999) University of California Santa Cruz; Ph.D. (2006) Brown University. Research specialties: marine ecology; marine conservation; coastal ecosystems; human impacts; environmental stress. Contact: andrewaltieri808@gmail.com

Baer, Boris, Research Associate. Biology (1993), M. S. (1995), Ph.D. (2000). ETH Zurich. Current Affiliation: University of California, Riverside. Research specialties: Sociobiology, Evolutionary biology, Sexual selection, Proteomics. Contact: boris.baer@uwa.edu.au

Barrios, Hector, Professor of Entomology, University of Panama. Ph.D. Instituto de Zoologia de la Academia de Sciencias. Research specialties: Entomology. Contact: BarriosH@si.edu

Batterman, Sarah, Research Associate. B. A. (2006), Ph.D. (2013). Current affiliation: University of Leeds. Research specialties:  Empirical, experimental and theoretical approaches of all levels of organization, from traits of individual plants to communities comprised of diverse species and plant functional types to emergent ecosystem and biome-level properties. Contact: s.a.batterman@leeds.ac.uk

Bernal, Ximena, Research Associate. B. S. (1999), M. S. (2001), Ph.D. (2007). Current Affiliation: Purdue University. Research specialties: Evolutionary Ecology, Animal Behavior, Communication. Contact: xbernal@purdue.edu

Bohlman, Stephanie, Research Associate. B. S. (1991), M. S. (1995), Ph.D. (2004). Current Affiliation: University of Florida. Research specialties: Forest Ecology, How species/functional group composition and forest structure will respond to climate change and the effects of these responses on ecosystem functioning. Contact: sbohlman@ufl.edu

Brawn, Jeffrey, Research Associate. B.S. (1975) University of Massachusetts, Amherst; M.S. (1979) University of Missouri, Columbia; Ph.D (1986) Northern Arizona University. Research specialties: Avian Ecology, tropical ecology, comparative life histories, demography. Contact: jbrawn@illinois.edu

Breedy, Odalisca, Research Associate. B.A. (1979), M.Sc. (1986), M.A.(2003) Universidad de Costa Rica. Research specialties: study of igneous and volcano-sedimentary processes and their link to the tectonic evolution of oceans and subduction zones.. Contact: odaliscab@gmail.com

Buchs, David, Research Associate. Ph.D. (2008) University of Lausanne. Research specialties: Octocoral taxonomy and biogeography, eastern Pacific and Caribbean. Contact:  dbuchs@cardiff.ac.uk

Chaverri, Gloriana, Research AssociateB.S. (1995) Universidad Nacional de Costa Rica; Ph.D. (2006) Boston University. Research specialties: Behavioral ecologist who focuses primarily on social behavior and sensory ecology. Contact:  gloriana.chaverri@ucr.ac.cr

Clay Keith, Research Associate.  B.S. (1977) Rutgers College; Ph.D (1982) Duke University.  Research Specialties: Species interactions, community ecology, biodiversity, disease, symbiosis. Contact: clay@indiana.edu

Coley, Phyllis, Research Affiliate. PhD (1981) University of Chicago. Research specialties: Tropical plant ecology; plant herbivore interactions, chemical ecology. Contact: coley@biology.utah.edu

Collins, Laurel, Research Associate. B.S. (1974) University of Maryland; M.S. (1985) George Washington University Ph.D. (1988) Yale University. Research area: The foraminifer assemblages in Holocene. Contact: collinsl@fiu.edu

Comita, Liza, Research Associate. B.A. (1999), M.A. (1999) University of Pennsylvania; Ph.D. (2006) University of Georgia. Research specialties: tropical forest ecology. Contact: liza.comita@yale.edu

Crofoot, Margaret, Research Associate. B.A. (2001) Stanford University; M.A. (2003), Ph.D. (2008) Harvard University. Research specialties: primate behavioral ecology, movement ecology. Contact: CrofootM@si.edu

Cusack, Daniela, Research Associate.   B.A. (1999) Wesleyan University;  M.Sc. (2003) Yale University; Ph. D. (2009) University of California – Berkeley, CA.  Research area: ecosystem ecology, biogeochemistry, and biophysical geography. Contact: dcusack@geog.ucla.edu

Dalling, James, Research Associate. B.A. (1988) Oxford University; Ph.D. (1992) Cambridge University. Research specialties: Seed ecology and forest regeneration. Contact: dallingj@life.illinois.edu

Dechman, Dina, Research Associate.  M. Sc. (1999), Ph.D. (2005). Current Affiliation: Max Plank Institute of Ornithology. Research specialties: Behavioural Ecology, Evolution. Contact: ddechmann@orn.mpg.de

De Leon, Luis Fernando, Research Associate. Ph. D. (2011) McGill University. Research Interests: Evolutionary ecology, adaptive divergence, ecological speciation, genomic of adaptation, gene flow, human impacts on evolution, eco-evolutionary dynamics Contact: luis.deleonreyna@umb.edu

Della Togna, Gina, Research Associate. Ph.D (2015). Current Affiliation: Universidad Interamericana de Panama. Research specialties: The reproductive physiology of amphibians for the development and application of Assisted Reproductive Technologies in endangered populations. gina.dellatogna@uip.pa

Dent, Daisy, Research Associate.  (1998),  Ph.D. (2004). Current Affiliation: University of Stirling. Research specialties: Tropical Ecology; identify the effects of human disturbance on tropical forest ecosystems. Contact: daisy.h.dent@gmail.com

Dick, Christopher William, Research Associate. B.A. (1990) Hampshire College; Ph.D. (1999) Harvard University. Research specialties: tropical forest ecology, biogeography, and ecological genetics of tropical forest trees. Contact: cwdick@umich.edu

Dudley, Robert, Research Associate. Ph.D. (1987) University of Cambridge. Research interest:  Biomechanics, functional morphology, and evolution of animal flight; Physiological ecology of tropical insects, particularly flight energetics and migration; Evolutionary origins of human alcoholism in primate frugivory.  Contact: wings@berkeley.edu

Engelbrecht, Bettina, Research Associate.  Ph.D. (2006) University of Darmstadt. Research interest: Functional Ecology, Ecophysiology, Tropical Ecology, Plant Population Ecology, Community Ecology, Global Change Ecology. Contact: bettina.engelbrecht@gmail.com

Escobar, Jaime, Research Associate.  BS (2002), M. Sc. (2005), Ph.D. (2010). Current Affiliation: Universidad del Norte. Research specialties: Geology, Natural resources. Contact: jhescobar@uninorte.edu.co

Fernandez, Hermogenes, Research Associate.  BS (1995), MS (2000), Ph.D. (2006). Current Affiliation:  INDICASAT. Research specialties: Evolution, Ecology, Behavior or leaf cutter ants, Host-parasite interactions, Diverisity and behaviour of honey bees. Contact: HFernandez@indicasat.org.pa

Freestone, Amy, Research Associate.  BS (2000), Ph.D. (2005). Current Affiliation: Temple University. Research specialties: Ecology and Conservation of Species Diversity Contact: amy.freestone@temple.edu

Gilbert, Gregory, Research Associate. B.S. (1985) State University of New York; M.S. (1988), Ph.D. (1991) University of Wisconsin, Madison. Current Affiliation: University of California, Santa Cruz. Research specialties: Plant disease ecology, fungal community ecology, and tropical forest ecology and conservation. Contact: ggilbert@ucsc.edu

Herrera, Amalia, Research Associate.  BS (1978), MS (1981), Ph.D. (1985). Research specialties: Taxonomy, Contact: herreraa@si.edu

Hiller, Alexandra, Research Associate. Ph.D. (2005) Justus-Liebig-University.  Research Interest: The evolutionary history of Porcellanidae. Contact: hillerh@si.edu

Holtum, Joseph, Research Associate.  Ph.D. (1980). Current Affiliation: James Cook University. Research specialties: Carbon metabolism in CAM plants and tropical epiphytes, Responses of tropical plants to increasing atmospheric CO2, Physiology and ecology of Australian desert plants, Ecophysiology of Australian ant-plants, Biofuel feedstocks for the wet-dry tropics, Responses of tropical biomes to drought Contact: joseph.holtum@jcu.edu.au

Jiggins, Christopher, Research Associate. B.A. Hons (1993) University of Cambridge; Ph.D. (1997) University College London. Current Affiliation: University of Cambridge. Research specialties: Genetic basis of adaptation and speciation in mimetic butterflies. Contact: c.jiggins@zoo.cam.ac.uk

Jones, Frank (Andy), Research Associate. Ph.D. (2004). Current Affiliation: Oregon State University. Research specialties: The Ecological and evolutionary mechanisms responsible for the origin and maintenance of plant diversity. Contact: jonesfr@science.oregonstate.edu

Kapheim, Karen, Research Associate.  Ph.D. (2010). Current affiliation: Utah State University. Research specialties: The evolutionary processes responsible for the diversity and plasticity of complex traits. Contact: karen.kapheim@usu.edu

Kays, Roland, Research Associate. B.A. (1993) Cornell University; Ph.D.(1999) University of Tennessee. Current Affiliation: North Carolina State University. Research specialties: Temperate and neotropical mammals, especially carnivores; radio-telemetry; non- invasive surveys; spatial ecology; canopy biology. Contact: rokays@gmail.com

Keiner, Christina, Research Associate. Ph.D. (2001), Johns Hopkins University; B.A. Biology (1993) McDaniel College. Research interest: The history of tropical environmental science and environmental politics. Contact: cmkgsh@rit.edu

Kitajima, Kaoru, Research Associate. B.A. (1984) University of Tokyo; M.S. (1987), Ph.D. (1992) University of Illinois, Champaign-Urbana. Current Affiliation: Kyoto University. Research specialties: Plant functional traits; physiological ecology of tropical trees; seed and seedling ecology; plant-soil interactions; functional basis for plant invasion. Contact: kaoruk@kais.kyoto-u.ac.jp

Knorschild, Mirjam, Research Associate. Biology (2005),  (2009), Ph.D. (2013). Ulm University.  Current Affiliation: Free University of Berlin. Research specialties: Animal Behavior, acoustic communication, cognition, learning and social behavior of mammals, particularly bats Contact: mirjam.knoernschild@gmail.com

Lasso, Eloisa, Research Associate. Biology (1996), M. Sc. (2001), Ph.D. (2007). Current Affiliation: University of Los Andes. Research specialties: Ecology y Plant physiology in the tropics. Contact: elolasso@gmail.com

Lasso,  Marixa, Research Associate. Ph.D (2002) University of Florida; M.A. (1996) University of Pittsburgh. Research Interest: Anthropology. Contact: mlasso@unal.edu.co

Leung, Brian, Research Associate. Ph.D. (1999) Carleton University; B.Sc (1993) University of British Columbia. Research Interest: Research and Integrated Sustainability Model Contact: brian.leung2@mcgill.ca

Linares, Mauricio, Research Associate. Biology (1983), Ph.D. (1989). University of Los Andes. Research specialties: Genética y Evolución Contact: mlinares@uniandes.edu.co

Loaiza, Jose, Research Associate. Biology (), M. S. (2004), Ph.D. (2010). Current Affiliation: INDICASAT. Research specialties: Disease ecologist with expertise in vector-borne pathogens. Contact: jose.loaiza@mail.mcgill.ca

Lopez, Omar, Research Associate. B. Sc. (1994), Ph.D. (2002). Current Affiliation: INDICASAT. Research specialties: Ecology and Evolution Contact: prioria@yahoo.com

Lucas, Peter, Research Associate. B. Sc. (1976), Ph.D. (1980). Current Affiliation: Kuwait University. Research specialties: Biological anthropology studying primate feeding ecology and feeding processes. Contact: peterwlucas@gmail.com

Mangan, Scott, Research Associate.  Ph.D (2008) Indiana University. Research Interest: Identifying the mechanisms that lead to the organization and function of biological communities is a fundamental goal in ecology.  Contact: smangan@wustl.edu

Mendizabal, Tomas, Research Associate. Ph.D. (2005) University College London; B.A. (1995) University of Liverpool. Research Interest: The later Precolumbian period in the eastern region of the country (AD 1 to 1500) and historical archaeology encompassing the Colonial, Departmental and Republican periods of Panamanian history from 1501 to the mid-20th century. Contact: tomas.mendizabal@gmail.com

Mejia, Luis, Research Associate. B. S. (2002),  Ph.D. (2009). Current Affiliation: INDICASAT. Research specialties: Plant-Microbe interactions; Biology, ecology, and systematics of plant associated fungi; Molecular Phylogenetics, Plant immunity. Contact: MejiaLC@si.edu

Merrill, Richard, Research Associate. M.Sc (2006) University of Oxford; Ph.D. University of Cambridge.  Research Interest: How the evolution and genetics of behaviour impact on animal diversity Contact: merrill@bio.lmu.de

Meylan, Peter, Research Associate. B.S. (1976), M.S. (1980), Ph.D. (1985) University of Florida. Current Affiliation: Florida Fish and Wildlife, Conservation Commission’s, Florida Marine Research Institute. Research specialties: Ecology, phylogeny, systematics and paleontology of reptiles especially turtles. Contact: Anne.Meylan@MyFWC.com

Meylan, Anne, Research Associate. B.S. (1974), M.S. (1978), Ph.D. (1984), University of Florida. Current Affiliation: Eckerd College. Research specialties: Reproductive biology, ecology and migrations of marine turtles; global status of hawksbill turtle (Eretmochelys imbricata). Contact: meylanpa@eckerd.edu

Moreno, Ricardo, Research Associate.  MS (2006). University of Panama. Research specialties: Population ecology.  Contact: rmoreno@yaguara.org

Ogden, Fred L., Research Associate. B.S. (1987), M.S. (1989), Ph.D. (1992) Colorado State University.  Current Affiliation: University of Wyoming. Research specialties: Hydrologic Model Development, Physical Hydrology, Hydrologic and Atmospheric Remote Sensing, Hydraulics, Ecohydrology. Contact: fogden@uwyo.edu

Parker, Ingrid, Research Associate. Ph.D. (1996). Current Affiliation: University of California, Santa Cruz. Research specialties: Population biologist, intersection of ecology and evolution, species interactions, particularly plant disease, pollination, and plant-human interactions  Contact: parker@biology.ucsc.edu

Potvin, Catherine, Research Associate. B. Sc. (), M. Sc. (), Ph.D. (1985). Current Affiliation: McGill University. Research specialties: Tropical forest ecology, conservation and climate change. The role of Indigenous peoples in climate change innovations. Contact: catherine.potvin@mcgill.ca

Puebla, Oscar, Research Associate. B. Sc. (2000), M. Sc. (2003), Ph.D. (2009). Current Affiliation: GEOMAR Helmholtz Centre for Ocean Research Kiel. Research specialties: Marine speciation, local adaptation, sexual selection, recombination, Genomics, Behavior. Contact: oscar.puebla@mail.mcgill.ca

Ranere, Anthony, Research Associate. B.A. (1964) Harvard University; M.A. (1968 Idaho State University; Ph.D. (1972) University of California, Davis. Research specialties: Archaeology of the American tropics; origins of agriculture; early human colonization of the Americas; lithic analysis. Contact: ranere@temple.edu

Ryan, Michael, Research Associate. B.A. (1975) Glassboro State College; M.S.(1977) Rutgers University; Ph.D. (1982) Cornell University. Current Affiliation: University of Texas, Austin. Research specialties: Evolution and mechanisms of animal behavior, especially animal communication and sexual selection. Contact: mryan@mail.utexas.edu

Sanchez-Azofeifa, G. Arturo, Research Associate. B.Sc. (1990) University of Costa Rica; M.Sc. (1993), Ph.D. (1996) University of New Hampshire. Current Affiliation: University of Alberta. Research specialties: Hyperspectral remote sensing, tropical dry forests, wireless sensor networks. Contact: arturo.sanchez@ualberta.ca

Santiago, Louis, Research Associate. Ph.D. (2003). Current Affiliation: University of California, Riverside. Research specialties: Environmental Physiology; Ecosystem Science; Evolutionary Physiology; Stable Isotopes; Tropical Biology. Contact: santiago@ucr.edu

Sayer, Emma, Research Associate. Ph.D. (2005). Current Affiliation: Lancaster University. Research specialties: Ecosystems response to change. Contact: e.sayer@lancaster.ac.uk

SCHNITZER, Stefan A., Research Associate. Ph.D. (2001) University of Pittsburgh. Current Affiliation: Marquette University. Research specialties: Plant Community Ecology; Tropical Forest Ecology; Plant Competition; Ecology of Lianas. Contact: S1@Marquette.edu

Schlinger, Barney, Research Associate. Ph.D. (1998). Current Affiliation: University of California, Los Angeles. Research specialties:  Contact: schlinge@lifesci.ucla.edu

Spalding, Ana, Research Associate. B. A. (1999), M. A. (2004), Ph.D. (2011). Current Affiliation: Oregon State University. Research specialties: Integrated coastal management, resource use and property regimes, international marine policy, and marine protected areas. Contact: ana.spalding@oregonstate.edu

Srygley, Robert, Research Associate. B.A. (1983) University of Washington; Ph.D. (1991) University of Texas. Current Affiliation: USDA. Research specialties: Physiology, behavior, evolution and ecology of insect migration, predator- prey interactions, and flight. Contact: robert.srygley@ars.usda.gov

Tanner, Edmund, Research Associate. B. Sc. (1973) University of London; Ph D (1977) University of Cambridge. Research specialties: Tropical forests, nutrients, water, trees, biodiversity, forest dynamics. ed.tanner@plantsci.cam.ac.uk

Tarwater, Corey, Research Associate. Ph.D. (2010) University of Illinois, Urbana-Champaign; M.S. (2006) University of Illinois, Urbana-Champaign; B.S. (1999) University of California, Davis. Research Interest: Temporal variation and impacts of climate change on tropical avian demography and avian communities and the role of the juvenile period in life history strategies and demography. Contact:  corey.tarwater@uwyo.edu

Taylor, Ryan, Research Associate. Ph.D. (2004). Current Affiliation: Salisbury University. Research specialties: Animal behavior; Role of multimodal signal assessment in sexual selection. Contact: rctaylor@salisbury.edu

Ter Hofstede, Hannah, Research Associate. Ph.D. (2008) University of Toronto; M.Sc. (2003) York University; B.Sc. (1998) University of Guelph. Research Interest: Sensory systems and how this process interacts with the behaviour and ecology of animals. Contact: hannah.th@gmail.com

Thacker Robert, Research Associate.  Ph.D. (1995) University of Michigan, Ann Arbor;  M.S. (1992) University of Michigan, Ann Arbor; B.S. (1990) Duke University.  Research Interest:  Understanding the ecology, evolution of sponge-microbe interactions and building new digital infrastructure for studying phenotype evolution. Contact: robert.thacker@stonybrook.edu

Tschapka, Marco, Research Associate. Ph.D. (2008). Current Affiliation: University of Ulm . Research specialties: Animal - plant interactions and Ecological background of emerging diseases Contact: marco.tschapka@uni-ulm.de

Valencia, Renato, Research Associate. Ph.D. (1994). Pontificia Universidad Catolicia del Ecuador. Research specialties: Evolution of vascular plants, tropical botany and forest ecology Contact: lrvalencia@puce.edu.ec

Van Bael, Sunshine, Research Associate. Ph.D. (2003). Current Affiliation: Tulane University. Research specialties: Community ecology, vertebrate ecology, microbial ecology, plant-animal-fungal interactions, multi-trophic interactions, tropical agriculture and biodiversity.. Contact: svanbael@tulane.edu

Van Breugel, Michiel, Research Associate.  MSc (1997), Ph.D. (2007). Current Affiliation: Yale NUS College. Research specialties: Ecology of forest and trees in human-modified landscapes Contact: mvbreugel@gmail.com

Velasquez-Runk, Julie, Research Associate. Ph.D. (2005). Current Affiliation: The University of Georgia. Research specialties: Cultural Anthropology. Contact: julievr@uga.edu

Villarreal A., Juan Carlos, Research Associate. Associate professor at Laval University, Quebec, Canada. B.Sc. (2003) Universidad de Panamá, M.Sc. (2005) Southern Illinois University, Carbondale, Ph.D. (2011) University of Connecticut, Postdoctoral (2012-2014) Botanischer Garten, München (LMU), Germany; (2015) Royal Botanic Garden, Edinburgh, Scotland; (2015) Smithsonian Tropical Research Institute.  Research specialties: Evolution, bryophyte taxonomy and systematics, symbiosis of plants (especially bryophytes and cycads), lichens and microbes. Contact: juan-carlos.villarreal-aguilar@bio.ulaval.ca. Site web: http://villarreal-lab.ibis.ulaval.ca/

Vollmer, Steven, Research Associate. B.Sc. (1994) Colorado State University; M.Sc. (1999) California State University Ph.D. (2004) Harvard University. Research Interest: Coral speciation and hybridization, the genetics coral immunity and disease resistance, and the nature and structure of coral microbiomes. Contact:  s.vollmer@northeastern.edu

Wake, Thomas, Research Associate.  Ph.D., (1995) University of California, Berkeley; M.A., (1988) University of California, Berkeley; B.A., (1986) University of California, Berkeley.  1986.  Research Interest: Archaeological and Zooarchaeological, development of Social Complexity; Neotropical Archaeology; Historical Archaeology; Paleoenvironmental Reconstruction; Human Impacts; Osteology. Contact: twake@ucla.edu

Wardle, David, Research Associate.  Ph.D (1989) University of Calgary. Research Interest: Understand linkages between the aboveground and belowground biota in southeast Asian rainforests and the consequences of these linkages for the functioning of these systems. Contact: David.Wardle@slu.se

Warkentin, Karen, Research Associate. B.Sc. (1985) University of Guelph; M.Sc. (1990) Dalhousie University; Ph.D. (1998) University of Texas, Austin. Current Affiliation: Boston University. Research specialties: Phenotypic plasticity, behavioral ecology & ecological developmental biology of amphibians, especially egg and larval stages; Predator- prey interactions and inducible hatching; Vibrations as an information source; Ecology of fear. Contact: KWarken@bu.edu

Wulf, Janie, Research Associate.  Ph.D. (1986). Current Affiliation: Florida State University. Research specialties: The ecology of sponges and the organisms with which they interact as mutualistic partners, competitors, and prey, especially in coral reefs, seagrass meadows, and mangroves. Contact: wulff@bio.fsu.edu

Yanoviak, Stephen, Research Associate.  Ph.D. (1990). Current Affiliation: University of Louisville. Research specialties: Tropical arthropod ecology, behavior and conservation. Contact: steve.yanoviak@louisville.edu

Zotz, Gerhard, Research Associate. M.Sc. (1989); Ph.D. (1993) University of Wuerzburg, Germany. Research specialties: Ecology of vascular and non-vascular epiphytes; tropical biodiversity; ecophysiology of mosses. Contact: Zotz@si.edu

 

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